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DNA research of all six dogs is presented below with for each microsatellite the number of the two alleles established :

 Marjolein has for microsatillite 1 (AHT 121) allele 102 in twofold. This allele is present in Lianne as well in Borus. For microsatellite 2 (AHT 137) Marjolein has the alleles 149 and 151. Allele 151 can be traced back to her dam Lianne, but her sire Borus (and her dam Lianne!) does not have allele 149! For microsatellite 3 (AHTH 171) Marjolein has the alleles 219 en 225, similar as her dam Lianne, but her sire Borus has the alleles 212 and 233. When you continue to check in this way all alleles for the 18 microsatellites present in the six animals, you can conclude that Borus cannot be the sire of Marjolein on base of the microsatellites 2, 3, 5, 6, 7, 13. On base of the alleles of Bart for the 18 microsatellites it may be very reasonable that Bart is the sire of Marjolein.

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A comparison of the alleles of Marjolein with the alleles of Lianne and of Bart indicate that Marjolein may be born out of a mating of Lianne and Bart and that Martha is born out of a mating of Lieneke and Boris. Apparently, something has been going wrong during the matings.